Ecosystem types

Estuarine realm

Lara van Niekerk1, 2 , Janine B. Adams3, 2 , Nicola C. James4, 2 , Stephen J. Lamberth5, 2 , Fiona C. Mackay6, 7 , Jane K. Turpie8 , Anusha Rajkaran9 , Steven P. Weerts1, 10 , Alan K. Whitfield4

1. Council for Scientific and Industrial Research

2. Institute for Coastal and Marine Research, Nelson Mandela University

3. DST/NRF Research Chair in Shallow Water Ecosystems, Nelson Mandela University

4. South African Institute for Aquatic Biodiversity

5. Department of Environment, Forestry and Fisheries

6. Oceanographic Research Institute, SAAMBR

7. University of KwaZulu-Natal

8. Anchor Environmental Consultants

9. University of the Western Cape

10. University of Zululand

Published

December 5, 2025

South Africa has 290 estuaries and 42 micro-estuaries, which have been classified into 22 estuarine ecosystem types and 3 micro-estuary types. This high diversity of estuary types stems from the country’s diverse climatic, oceanographic and geological drivers.

The Goukamma Estuary, a Warm Temperate Large Temporarily Closed type, is in a Marine Protected Area and in a near-natural condition, yet under pressure from flow reduction, pollution, fishing, riparian development and artificial breaching. (© Cape Nature)

Estuary ecosystem types can serve as surrogates for ecosystem processes and enable predictions of biophysical characteristics. Understanding ecological processes and patterns associated with an estuary type facilitates an assessment of its resilience to anthropogenic pressures. This allows for extrapolation in data-limited environments (see Figure 1).

The estuarine classification scheme divided the biogeographical regions that characterise the South African coast into four major zones: the Cool Temperate (Orange to Ratel), the Warm Temperate (Heuningnes to Mendwana), the Subtropical (Mbashe to St Lucia) and the Tropical (uMgobezeleni to Kosi), with the latter being a new addition to the estuarine biogeographical regions.

All rivers or streams with outlets to the sea were categorised broadly as estuaries and micro-systems. South Africa’s 290 estuaries are categorised into nine functional types, namely Estuarine Lake, Estuarine Bay, Estuarine Lagoon (e.g. the groundwater-fed Langebaan system), Predominantly Open, Large and Small Temporarily Closed, Large and Small Fluvially Dominated, and Arid Predominantly Closed. Micro-systems were further divided into micro-estuaries (42), micro-outlets, and coastal waterfalls based on spatial features and biotic observations in the field.

Figure 1. Estuarine ecosystem typology

Biographical regions

The traditional biogeographic organisation of South African estuaries has included only three dominant regions, viz. subtropical, warm temperate and cool temperate13. However, the tropical distribution of some species recorded in the Kosi and uMgobezeleni estuaries in the north, prompted subdividing the subtropical region and include a tropical transition zone in the north-east of South Africa. Both of these systems, albeit differing in size and function, are unique clear water estuaries on a small section of South African coast.

Kosi Estuary has uniquely tropical flora indicated by presence of the seagrass (Halodule universis) and mangrove species Lumnitzera racemosa, Ceriops tagal and Xylocarpus granatum that are found in no other South African systems4. Tropical species such as the snake sea cucumber (Synapta maculate) and pen shell (Pinna muricata) (associated with H. universis) also point to a tropical influence on the system’s invertebrate fauna. The ichthyofaunal comprises a wide diversity of fishes, including tropical species not reported from any other South African estuaries5. This is evidenced by the abundance and large sizes of several members of the Lutjanidae (snappers) in the system.

St Lucia Estuary, to the south of Kosi, supports invertebrate communities common in subtropical estuaries6. The tropical zone, therefore, aligns with marine biogeographic patterns evident in the marine biogeographic map (the Natal-Delagoa tropical region)7.

Thus, the Estuarine Ecosystem Classification scheme defined the biogeographical regions as:

  • Cool temperate from Orange Estuary on the Northern Cape to Uilkraals Estuary in the Western Cape;

  • Warm temperate from the Ratel Estuary near Cape Agulhas to Mendwana Estuary in the Eastern Cape;

  • Subtropical stretches from the Mbashe Estuary in the Eastern Cape to the St Lucia Estuary in KwaZulu-Natal; and

  • Tropical from uMgobezeleni to Kosi Estuary in Northen KwaZulu-Natal.

What is an estuary?

The NBA 2025 defines an estuary as ‘a partially enclosed permanent water body, either continuously or periodically open to the sea on decadal time scales, extending as far as the upper limit of tidal action, salinity penetration or back-flooding under closed mouth conditions. During floods an estuary can become a river mouth with no seawater entering the formerly estuarine area or, when there is little or no fluvial input, an estuary can be isolated from the sea by a sandbar and become fresh or even hypersaline4. A defining feature of this definition is that complex estuarine abiotic processes distinguish estuaries from other aquatic ecosystem types, i.e. restricted tidal action, mixing of fresh and salt water, increased retention and/or increased water levels under closed mouth conditions.

Estuary functional types

South Africa’s 290 estuaries are categorised into nine functional types, namely Estuarine Lake, Estuarine Bay, Estuarine Lagoons (e.g. the groundwater-fed Langebaan system), Predominantly Open, Large and Small Temporarily Closed, Large and Small Fluvially Dominated, and Arid Predominantly Closed (see Figure 2).

Figure 2. Examples of South African estuary types.

Estuarine Lakes

Estuarine Lakes comprise one or more typically large circular water bodies connected to the sea by a constricted inlet channel. Maximum water levels are determined by berm height, mouth state and freshwater input. Marine connectivity varies from almost permanently open to closed on annual scales. Salinities are highly variable, ranging from fresh to hypersaline because of differing freshwater input (surface and ground water), evaporation and the extent of the marine connection. Mixing processes are dominated by wind and, to a lesser extent fluvial inputs, owing to their restricted mouths and relatively large surface areas. Sediment processes tend to be stable, with infilling occurring over long time scales and resetting confined to larger flood events.

Estuarine Bays

Estuarine Bays are permanently linked to the sea by unrestricted, deep mouths and are dominated by tidal processes, with tidal amplitudes close to those of the sea. These are large systems (>1 200 ha) with generally round basins where only the upper reaches experience a degree of constriction to tidal flows. As a result of relatively low river inputs they have a predominantly euhaline salinity regime in the lower and mid reaches, with freshwater mixing processes being mostly confined to the restricted upper areas. Sediments are typically marine in origin and grain size distributions are stable. There are two natural occurrences of Estuarine Bays in South Africa (viz. Knysna and Durban Bay).

Estuarine Lagoons

Langebaan Lagoon has many of the characteristics of an estuary8, including calm waters that are protected from marine wave action and biota that reflect many of the species usually found in estuaries. However, despite groundwater seeps into some areas, it lacks surface water inflow and a normal estuarine salinity gradient. Langebaan Lagoon represents a unique coastal ecosystem type and is recognised as an estuary because its ecological functioning includes both freshwater and marine inputs into a semi-enclosed embayment. Estuarine Lagoons, as defined here, are permanently connected to the sea and are therefore marine dominated. Tidal action is the dominant mixing process and sedimentary processes are thus generally stable. Tidal amplitude and water levels are close to those of the sea.

Predominantly Open estuaries

These estuaries are open to the sea for more than 90% of the time. Some are permanently open owing to perennial river flows or the presence of large tidal prisms. Predominantly Open estuaries are linear systems in which mixing processes are dominated by both fluvial inputs and tidal action creating vertical and horizontal salinity gradients. Under low flows, hypersalinity can develop in the upper reaches. The degree to which the mouth is restricted depends on the rate and volume of freshwater inflow. Some systems become severely constricted during low flow periods. Regular flooding results in relatively mobile sediments. These estuaries usually support wetlands, salt marshes, macrophyte beds and marine and estuarine fauna. Surprisingly, their size varies considerably ranging from 10 to 7500 ha, with smaller systems afforded a degree of protection against direct wave action by rocky headlands or subtidal reefs, which assists in maintaining an open mouth.

Large and Small Temporarily Closed estuaries

The sizeable temporarily open/closed category of Whitfield (1992)9 was divided into Small and Large Temporarily Closed estuaries using a total habitat area of 15ha (associated with ~10 ha of open water area) as the dividing threshold. The division was based on differences in biophysical processes and patterns. Small systems are likely to experience rapid increases and decreases in freshwater runoff over a few hours making them strongly event driven. There will be little scouring during breachings, and a semi-closed mouth condition can easily develop owing to a small, perched, outflow channel that restricts tidal amplitude to 10 to 15 cm. There is minimal water column area during the open state. Habitat diversity is likely to be low (and without intertidal salt marshes) and species diversity and abundance reduced.

Both large and small systems tend to be linear or funnel shaped with highly restricted inlets. Smaller systems especially tend to be ‘perched’ above normal tidal levels, resulting in little to no open water area during the open mouth low tide state. Water levels are dominated by the state of the mouth, with highest levels of 1 m to 2 m above mean sea level during the closed phase. Open phase mixing processes are dominated by fluvial input and partially by tides. When closed, wind and seepage losses through the berm play a key role. Sediment composition is largely stable, resetting mainly during floods. Salinity regimes range from almost fresh to hypersaline, as that latter can develop during times of low flow or droughts in large systems. Small Temporarily Closed estuaries tend to be fresher in character as they have less connectivity with the sea.

Small and Large Fluvially Dominated estuaries

The classification distinguishes between small (<15 ha), black water dominated, rocky temperate southern coast estuaries; and large (≥15 ha), shallow, sediment rich, freshwater dominated systems of the east and west coasts. The larger systems have very high sediment turnover, often develop ebb-tidal deltas, are turbid and can close during periods of low flow, e.g. uThukela and Orange estuaries. Small, sediment-starved, fluvially-dominated systems have unrestricted mouths as they usually occur along rocky shores and receive clear humic-stained water from Table Mountain Sandstone catchments. Large Fluvially Dominated estuaries tend to be constricted and can even periodically close during low flows. River processes are dominant and salinities are mostly fresh throughout the estuary for more than half the time. During peak flood conditions, outflows can influence salinities for considerable distances offshore.

Arid Predominantly Closed estuaries

This type comprises six small estuaries (Buffels, Swartlintjies, Spoeg, Groen, Sout (Noord) and Wadrift) located in the arid Namaqua West Coast region. They are linear or funnel shaped and closed on annual to decadal scales. Salinities tend to be high to hypersaline (>35) as a result of low fluvial input and high evaporation rates. Thus, mixing processes tend to occur over long time periods and are dominated by the effects of evaporation, winds and seepage. Occasional breaching and over-washing during high sea conditions provide for marine connectivity. Sediment processes are generally stable on decadal time scales and are reset by large intermittent flash floods. Water levels are determined by the interplay between sand berm level, evaporation rates and seepage losses. Groundwater and inflows from local fountains replenish these losses and influence the salinity regimes of these estuaries.

Arid conditions promote the growth of unique vegetation such as salt tolerant, succulent Sarcocornia spp. and Salicornia spp. that can flourish despite open water salinities reaching above 200. Fish species rely on ‘suicidal’ recruitment that is largely a function of connectivity with the sea and overwash during high seas. Fish survival depends on groundwater maintaining salinity gradients and hypersalinity not exceeding 40 in parts of the systems. Invertebrate diversity, abundance and community structure are related to changing salinity gradients including long cycles of hypersalinity, e.g. Shrimp Artemia sp. that hatch at salinities above 40 and encyst when salinities exceed 150. Cycles of Artemia abundance follow salinity regimes that in turn affect the diversity, abundance and occurrence of flamingos and other birds that feed on them.

Estuarine ecosystem types in South Africa

The Estuarine Ecosystem Classification system for South Africa’s 290 estuaries are derived from interplay between biogeographical regions and estuary functional types, stemming from the country’s diverse climatic, oceanographic and geological drivers.

Estuarine Lagoons are the rarest South African estuary type with only one member in the Cool Temperate region, followed by Estuarine Bays with two in the Subtropical and one in the Warm Temperate region. Arid Predominantly Closed estuaries are confined to six systems in the Cool Temperate region. The Large and Small Fluvially Dominated types comprise seven systems each, occurring in three and two biogeographical regions, respectively. Small Temporarily Closed (116), Large Temporarily Closed (93), and Predominantly Open (45) are the most dominant types occurring across the Cool Temperate, Warm Temperate and Subtropical biogeographical regions. Estuarine Lakes occur across all four biogeographical zones. While not numerically dominant, this type represents the largest surface area of all estuary functional types, with Lake St Lucia representing more than half of South Africa’s estuarine surface area.

Estuary name Estuary type Estuary functional type Protection level Threat status Condition Extent (ha)
Orange Cool Temperate Large Fluvially Dominated Cool Temperate Poorly Protected Endangered Heavily modified 3019.1
Buffels Cool Temperate Arid Predominantly Closed Cool Temperate Poorly Protected Endangered Heavily modified 87.6
Swartlintjies Cool Temperate Arid Predominantly Closed Cool Temperate Poorly Protected Endangered Near-natural 134.3
Spoeg Cool Temperate Arid Predominantly Closed Cool Temperate Poorly Protected Endangered Near-natural 88.2
Groen Cool Temperate Arid Predominantly Closed Cool Temperate Poorly Protected Endangered Near-natural 88.2
Sout (Noord) Cool Temperate Arid Predominantly Closed Cool Temperate Poorly Protected Endangered Severely / critically modified 532.8
Olifants Cool Temperate Predominantly Open Cool Temperate Not Protected Endangered Moderately modified 2778.6
Jakkals Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Heavily modified 72.6
Wadrift Cool Temperate Arid Predominantly Closed Cool Temperate Poorly Protected Endangered Heavily modified 509.0
Verlorenvlei Cool Temperate Estuarine Lake Cool Temperate Poorly Protected Endangered Severely / critically modified 2276.3
Groot Berg Cool Temperate Predominantly Open Cool Temperate Not Protected Endangered Heavily modified 11733.5
Langebaan Cool Temperate Estuarine Lagoon Cool Temperate Moderately Protected Endangered Near-natural 6006.9
Diep/Rietvlei Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Heavily modified 1250.0
Sout (Wes) Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Severely / critically modified 613.7
Disa Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Severely / critically modified 23.5
Wildevoëlvlei Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Heavily modified 604.8
Schuster Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Near-natural 22.6
Krom Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Natural 100.5
Silwermyn Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Severely / critically modified 15.8
Zand Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Heavily modified 381.8
Zeekoei Cool Temperate Estuarine Lake Cool Temperate Poorly Protected Endangered Severely / critically modified 405.5
Eerste Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Severely / critically modified 277.4
Lourens Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Heavily modified 266.5
Sir Lowry’s Pass Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Severely / critically modified 4.1
Steenbras Cool Temperate Small Fluvially Dominated Cool Temperate Well Protected Vulnerable Near-natural 4.0
Rooiels Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Natural 21.6
Buffels (Oos) Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Near-natural 35.8
Palmiet Cool Temperate Large Temporarily Closed Cool Temperate Poorly Protected Endangered Moderately modified 44.2
Bot/Kleinmond Cool Temperate Estuarine Lake Cool Temperate Poorly Protected Endangered Moderately modified 2345.9
Onrus Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Heavily modified 30.8
Klein Cool Temperate Estuarine Lake Cool Temperate Poorly Protected Endangered Moderately modified 2065.5
Uilkraals Cool Temperate Predominantly Open Cool Temperate Not Protected Endangered Heavily modified 747.7
Ratel Cool Temperate Small Temporarily Closed Cool Temperate Moderately Protected Vulnerable Near-natural 35.9
Heuningnes Warm Temperate Estuarine Lake Warm Temperate Moderately Protected Endangered Heavily modified 10993.7
Klipdrifsfontein Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 6.7
Breede Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 2257.6
Duiwenhoks Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 441.4
Goukou Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 438.7
Gouritz Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 1099.7
Blinde Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 19.9
Tweekuilen Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Severely / critically modified 23.2
Gericke Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Severely / critically modified 8.7
Hartenbos Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Heavily modified 306.1
Klein Brak Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Heavily modified 1021.7
Groot Brak Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Heavily modified 318.1
Maalgate Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 34.4
Gwaing Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 21.8
Kaaimans Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 40.6
Touw/Wilderness Warm Temperate Estuarine Lake Warm Temperate Moderately Protected Endangered Moderately modified 1180.4
Swartvlei Warm Temperate Estuarine Lake Warm Temperate Moderately Protected Endangered Near-natural 2168.2
Goukamma Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 299.2
Knysna Warm Temperate Estuarine Bay Warm Temperate Moderately Protected Vulnerable Near-natural 3010.6
Noetsie Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 50.6
Piesang Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Heavily modified 87.3
Keurbooms Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 1623.8
Matjies Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 12.1
Sout (Oos) Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Natural 24.1
Groot (Wes) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 100.0
Bloukrans Warm Temperate Small Fluvially Dominated Warm Temperate Well Protected Least Concern Natural 15.3
Lottering Warm Temperate Small Fluvially Dominated Warm Temperate Well Protected Least Concern Natural 11.2
Elandsbos Warm Temperate Small Fluvially Dominated Warm Temperate Well Protected Least Concern Natural 22.1
Storms Warm Temperate Small Fluvially Dominated Warm Temperate Well Protected Least Concern Natural 12.4
Elands Warm Temperate Small Fluvially Dominated Warm Temperate Well Protected Least Concern Near-natural 31.1
Groot (Oos) Warm Temperate Small Fluvially Dominated Warm Temperate Well Protected Least Concern Natural 30.3
Tsitsikamma Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 67.4
Klipdrif (Oos) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 66.8
Slang Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 42.8
Kromme Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Heavily modified 755.5
Seekoei Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Heavily modified 322.4
Kabeljous Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 311.9
Gamtoos Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Heavily modified 2133.4
Van Stadens Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 101.8
Maitland Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 67.6
Baakens Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Severely / critically modified 13.5
Papkuils Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Severely / critically modified 80.4
Swartkops Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Heavily modified 2861.0
Coega (Ngqurha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Severely / critically modified 443.5
Sundays Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 824.0
Boknes Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 47.5
Bushmans Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 888.8
Kariega Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 559.0
Grant's Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 16.3
Kasouga Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 247.6
Kowie Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 484.9
Rufane Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 24.7
Riet Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 40.3
West Kleinemonde Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 224.2
East Kleinemonde Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 100.9
Great Fish Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 519.9
Old Woman’s Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Moderately modified 23.3
Mpekweni Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 83.1
Mtati (Mthathi) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 142.1
Mgwalana Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 216.9
Bira (Bhirha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 270.6
Gqutywa Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 105.4
Ngculura (Ngculurha) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 36.6
Mtana Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 81.2
Keiskamma Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 1402.0
Nqinisa Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 54.3
Kiwane (Khiwane) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Natural 87.0
Tyolomnqa Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 207.6
Shelbertsstroom Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 15.0
Lilyvale Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 33.7
Ross' Creek Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 13.0
Ncera (Ncerha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 77.9
Mlele Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 20.2
Mcantsi Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 31.8
Gxulu Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Moderately modified 106.6
Goda Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 51.9
Hlozi Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 40.9
Hickman's Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Moderately modified 25.9
Buffalo Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Heavily modified 137.5
Blind Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Heavily modified 14.3
Hlaze (iHlanze) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Heavily modified 7.9
Nahoon Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Moderately modified 178.5
Qinira (Quinirha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 110.7
Gqunube Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 176.3
Kwelera (Kwelerha) Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 121.6
Bulura (Bulurha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 135.4
Cunge Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 8.7
Cintsa Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 122.6
Cefane Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 176.5
Kwenxura (Kwenxurha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Natural 126.1
Nyara (Nyarha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 55.3
Imtwendwe (Mtwendwe) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 15.9
Haga-haga Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 25.5
Mtendwe Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 17.1
Quko Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Natural 102.1
Morgan Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 83.5
Cwili Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 14.7
Great Kei Warm Temperate Large Fluvially Dominated Warm Temperate Not Protected Vulnerable Moderately modified 572.6
Gxara (Gxarha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 68.5
Ngogwane Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 22.5
Qolora (Qolorha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 69.3
Ncizele Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 22.9
Timba Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 9.2
Kobonqaba (Khobonqaba) Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 115.1
Nxaxo/Ngqusi Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 269.5
Cebe Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Natural 82.1
Gqunqe Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Natural 79.1
Zalu Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 68.8
Ngqwara (Ngqwarha) Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 78.9
Sihlontlweni Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 55.2
Nebelele Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 29.8
Qora (Qhorha) Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 133.9
Jujura (Jujurha) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 30.3
Ngadla Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 40.7
Shixini Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Natural 116.4
Beechamwood Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 20.1
Kwazlelitsha (Kwazwedala) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 32.1
Kwa-Goqo Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 30.9
Ku-Nocekedwa Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 33.7
Nqabara/Nqabarana Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Near-natural 307.8
Ngomane (East) Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Near-natural 25.9
Ngoma/Kobule Warm Temperate Small Temporarily Closed Warm Temperate Moderately Protected Least Concern Natural 88.8
Mendu Warm Temperate Large Temporarily Closed Warm Temperate Poorly Protected Vulnerable Near-natural 107.4
Mendwana Warm Temperate Predominantly Open Warm Temperate Poorly Protected Vulnerable Natural 27.8
Mbashe Subtropical Large Fluvially Dominated Subtropical Poorly Protected Endangered Moderately modified 469.8
Ku-Mpenzu Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 31.7
Ku-Bhula (Mbhanyana) Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 58.7
Kwa-Suku Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 21.4
Ntlonyane Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 89.7
Nkanya Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 62.1
Sundwana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 25.2
Xora Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 261.5
Bulungula Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 67.4
Ku-Amanzimuzama Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 13.0
Nqakanqa Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 16.9
Mdikana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 17.0
Mncwasa Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 116.7
Mpako Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 51.5
Nenga Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 33.6
Mapuzi Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 12.2
Mtata Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Moderately modified 545.6
Thsani Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 12.6
Mdumbi Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 254.9
Lwandilana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 49.6
Lwandile Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 100.5
Mtakatye Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 247.8
Hluleka Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 30.1
Mnenu Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 197.0
Mtonga Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 56.2
Mpande Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 51.3
Sinangwana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 123.1
Mngazana Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 770.6
Mngazi Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 434.3
Gxwaleni Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 16.1
Bulolo Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 13.2
Mtumbane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 15.5
Mzimvubu Subtropical Large Fluvially Dominated Subtropical Poorly Protected Endangered Near-natural 842.7
Ntlupeni Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 30.8
Nkodusweni Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 100.0
Mntafufu Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 277.5
Ingo Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 34.7
Mzintlava Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 108.9
Mzimpunzi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 67.8
Kwanyambalala Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 24.8
Mbotyi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 64.1
Mkozi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 38.1
Sikatsha Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 13.3
Lupatana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 14.5
Mkweni Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 15.4
Msikaba Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 142.0
Mgwegwe Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 19.4
Mgwetyana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 22.1
Mtentu Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 92.8
Sikombe Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 102.1
Kwanyana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 57.8
Mtolane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Natural 46.5
Mnyameni Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 87.5
Mpahlanyana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 24.1
Mpahlane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 35.2
Mzamba Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Near-natural 110.7
Mtentwana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 33.4
uMthavuna Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 160.0
iSolwane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 18.7
iSandlu Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 32.4
uMbhoyibhoyi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 20.2
uMuntongazi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 11.5
iKhandalendlovu Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 31.6
iMpenjani Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 100.8
uMhlangomkhulu (South) (Umhlangankulu) Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 45.5
iKhaba Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 26.2
iMbizana Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 95.9
iMvutshini Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 10.7
iBilanhlonhlo Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Heavily modified 23.6
uVuzana Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 9.8
iKongeni Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Severely / critically modified 44.8
uVunguza Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 14.5
oHlangeni Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 70.3
iZotsha Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 104.0
iBhobhoyi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 47.9
uMbango Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Severely / critically modified 36.7
uMzimkhulu Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Moderately modified 402.9
uMthente Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 102.5
uMhlangomkhulu (North) (Mhlangamkulu) Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 27.9
iDombe Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Heavily modified 27.1
iKhoshwana Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 23.1
iNjambili Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 67.4
uMzumbe Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 291.2
uMhlabashana Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Near-natural 32.5
uMhlungwa Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 43.8
uMfazezala Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 20.9
uMakhosi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 19.0
uMnamfu Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 18.1
uMthwalume Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 185.6
uMvuzi Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 36.7
iFafa Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Heavily modified 111.2
uMdesingane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Heavily modified 25.0
iSezela Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 72.7
uMkhumbane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 24.8
iNkomba Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 1.1
uMuziwezinto Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 105.3
uMzimayi Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 26.6
Rocky Bay Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Heavily modified 5.9
uMphambanyoni Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 84.4
aMahlongwa Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Moderately modified 98.1
uMahlongwane Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 40.7
uMkhomazi Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Moderately modified 331.9
iNgane Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Heavily modified 34.6
uMgababa Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 138.0
uMsimbazi Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 193.7
iLovu Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Heavily modified 425.4
aManzanamtoti Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Severely / critically modified 55.2
aManzimtoti Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Severely / critically modified 50.3
iZimbokodo Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Severely / critically modified 89.4
iSiphingo Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Severely / critically modified 704.4
Durban Bay Subtropical Estuarine Bay Subtropical Not Protected Critically Endangered Severely / critically modified 2815.2
uMngeni Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Severely / critically modified 403.5
uMhlanga Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Severely / critically modified 150.8
uMdloti Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Heavily modified 189.5
uThongathi Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Severely / critically modified 145.6
uMhlali Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Heavily modified 200.9
Bob’s Stream Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Heavily modified 8.9
uSetheni Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Near-natural 10.2
uMvoti Subtropical Large Fluvially Dominated Subtropical Poorly Protected Endangered Heavily modified 288.7
uMdlotane Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 79.9
iNonoti Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 153.8
iZinkwazi Subtropical Large Temporarily Closed Subtropical Poorly Protected Vulnerable Moderately modified 300.1
uThukela Subtropical Large Fluvially Dominated Subtropical Poorly Protected Endangered Heavily modified 1261.0
aMatigulu/iNyoni Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Moderately modified 1183.1
iSiyaya Subtropical Small Temporarily Closed Subtropical Moderately Protected Vulnerable Severely / critically modified 88.4
uMlalazi Subtropical Predominantly Open Subtropical Poorly Protected Vulnerable Moderately modified 1851.7
uMhlathuze Subtropical Estuarine Lake Subtropical Moderately Protected Endangered Heavily modified 5642.3
Richards Bay Subtropical Estuarine Lake Subtropical Moderately Protected Endangered Severely / critically modified 6365.1
iNhlabane Subtropical Estuarine Lake Subtropical Moderately Protected Endangered Severely / critically modified 1875.7
iMfolozi/uMsunduze Subtropical Large Fluvially Dominated Subtropical Poorly Protected Endangered Heavily modified 7903.7
St Lucia Subtropical Estuarine Lake Subtropical Moderately Protected Endangered Heavily modified 67992.2
uMgobezeleni Tropical Estuarine Lake Tropical Moderately Protected Vulnerable Near-natural 829.3
Kosi Tropical Estuarine Lake Tropical Moderately Protected Vulnerable Near-natural 7337.0

Download the data here.

Technical documentation

Data availability

  • Estuarine Ecosystem Map: BGIS

Key publications

Van Niekerk, L, Adams JB, James, N, Lamberth S, Mackay F, Rajkaran A, Turpie J, Weerts S, Whitfield AK. 2020. An Estuary Ecosystem Classification that encompasses biogeography and a high diversity of types in support of protection and management. African Journal of Aquatic Science, 45: 199-216. doi: https://doi.org/10.2989/16085914.2019.1685934

References

1. Emanuel, B. et al. 1992. A zoogeographic and functional approach to the selection of marine reserves on the west coast of south africa. South African Journal of Marine Science 12: 341354.
2. Harrison, T.D. 2002. Preliminary assessment of the biogeography of fishes in south african estuaries. Marine and Freshwater Research 53: 479–490.
3. Turpie, J.K. et al. 2000. Biogeography and the selection of priority areas for conservation of south african coastal fishes. Biological Conservation 92: 59–72. https://doi.org/10.1016/S0006-3207(99)00063-4
4. Van Niekerk, L. et al. 2020. An estuary ecosystem classification that encompasses biogeography and a high diversity of types in support of protection and management. African Journal of Aquatic Science 45: 199216.
5. Blaber, S. & C. DP. 1981. A revised checklist and further notes on the fishes of the kasi system.
6. MacKay, F. et al. 2010. Macrobenthic invertebrate responses to prolonged drought in south africa’s largest estuarine lake complex. Estuarine, Coastal and Shelf Science 86: 553567.
7. Sink, K.J. et al. 2019. National biodiversity assessment 2018: Technical report volume 4: Marine realm. South African National Biodiversity Institute, Pretoria.
8. Whitfield, A. 2005. Langebaana new type of estuary? African Journal of Aquatic Science 30: 207209.
9. Whitfield, A. 1992. A characterization of southern african estuarine systems. Southern African Journal of Aquatic Science 18: 89103.